Osmolyte production by abundant ammonia oxidising archaea: A novel link between marine nitrogen and sulfur cycling?

(LEHTOVIRTAMORLEY_UBIO24ARIES)

Osmolyte production by abundant ammonia oxidising archaea: A novel link between marine nitrogen and sulfur cycling?

(LEHTOVIRTAMORLEY_UBIO24ARIES)

Project Description

Supervisors

Dr Laura Lehtovirta-Morley, University of East Anglia – Contact me

Professor Jonathan Todd, School of Biological Sciences, University of East Anglia

Dr Matthew Wallace, School of Pharmacy, University of East Anglia

Scientific background

Ammonia oxidising archaea (AOA) perform a critical step in the global nitrogen cycle and thrive in oceans with massive populations that account for ~ 40% of marine prokaryotes. Most microorganisms produce compounds called osmolytes to cope with environmental stresses including high salinity and drought, but the metabolic pathways and environmental importance of osmolyte production by AOA remains vastly underexplored. In new pilot data we show that abundant AOA produce glycine betaine (GB) and dimethylsulfoniopropionate (DMSP), two ubiquitous marine osmolytes that are critical in global nitrogen and sulfur cycling, respectively, and are major sources of climate-active gases that can impact Earth’s climate. This project will elucidate the metabolic pathways and the significance of GB and DMSP production by AOA in Earth’s oceans. This project is exciting and novel because it explores the much-overlooked role of a hugely abundant group of marine microorganisms in linking nitrogen and sulfur cycling. Furthermore, the project will reveal insights into adaptation strategies of these ubiquitous microbes in the ocean. This project is timely due to the key roles of biogeochemical nitrogen and sulfur cycling in climate change and environmental change.

Research methodology

The student will use multi-omics (transcriptomics, proteomics and metabolomics) approaches with ammonia oxidising archaeal cultures to examine the effect of e.g., salinity and temperature, on osmolyte production and establish up- and downregulated metabolism. Genes responsible for osmolyte production will be cloned, expressed and functionally characterised in host bacteria mutated in varied osmolyte production. Environmental distribution and expression of novel osmolyte pathways will be examined using metagenomics and metatranscriptomics datasets from diverse environments including coastal ecosystems, sediments, deep ocean trenches and open oceans.

Training

The student will be trained in diverse molecular microbial ecology, including microbial cultivation and physiology, analytical chemistry, nucleotide extraction, cloning, protein expression and characterisation, and omics-based approaches including bioinformatics. The student will join a thriving cohort of PhD students at UEA, collaborate with colleagues in Germany and the Netherlands, and present their research in lab meetings, research seminars and national and international conferences.

Person specification

This project is suitable for candidates with a BSc in microbiology-related disciplines.

References

  • 1 Zhong H, Lehtovirta-Morley LE, Liu J, Zheng Y, Lin H, Song D, Todd JD, Tian J, Zhang XH. Novel insights into the Thaumarchaeota in the deepest oceans: their metabolism and potential adaptation mechanisms. Microbiome (2020) 8:78.
  • 2 Widderich N, Czech L, Elling FJ, Könneke M, Stöveken N, Pittelkow M, Riclea R, Dickschat JS, Heider J, Bremer E. Strangers in the archaeal world: osmostress-responsive biosynthesis of ectoine and hydroxyectoine by the marine thaumarchaeon Nitrosopumilus maritimus: ectoine and hydroxyectoine biosynthesis in Archaea. Environ Microbiol (2016) 18:1227–1248.Widderich N, Czech L, Elling FJ, Könneke M, Stöveken N, Pittelkow M, Riclea R, Dickschat JS, Heider J, Bremer E. Strangers in the archaeal world: osmostress-responsive biosynthesis of ectoine and hydroxyectoine by the marine thaumarchaeon Nitrosopumilus maritimus: ectoine and hydroxyectoine biosynthesis in Archaea. Environ Microbiol (2016) 18:1227–1248.
  • 3 Oudova-Rivera B, Wright CL, Crombie AT, Murrell JC, Lehtovirta-Morley LE. The effect of methane and methanol on the terrestrial ammonia-oxidizing archaeon 'Candidatus Nitrosocosmicus franklandus C13'. Environ Microbiol (2023) 25(5):948-961.
  • 4 Curson ARJ, Williams BT, Pinchbeck BJ, Sims LP, Martínez AB, Rivera PPL, Kumaresan D, Mercadé E, Spurgin LG, Carrión O, Moxon S, Cattolico RA, Kuzhiumparambil U, Guagliardo P, Clode PL, Raina JB, Todd JD. (2018). DSYB catalyses the key step of dimethylsulfoniopropionate biosynthesis in many phytoplankton. Nature Microbiology. 4: 430-439.
  • 5 Williams BT, Cowles K, Bermejo Martínez A, Curson ARJ, Zheng Y, Liu J, Newton-Payne S, Hind AJ, Li CY, Rivera PPL, Carrión O, Liu J, Spurgin LG, Brearley CA, Wagner Mackenzie B, Pinchbeck BJ, Peng M, Pratscher J, Zhang XH, Zhang YZ, Murrell JC, Todd JD. (2019). Nature Microbiology, 4 (11), 1815-1825.

Key Information

  • This project has been shortlisted for funding by the ARIES NERC DTP and will start on 1st October 2024. The closing date for applications is 23:59 on 10th January 2024.
  • Successful candidates who meet UKRI’s eligibility criteria will be awarded a NERC studentship, which covers fees, stipend (£18,622 p.a. for 2023/24) and research funding. International applicants are eligible for fully-funded ARIES studentships including fees. Please note however that ARIES funding does not cover additional costs associated with relocation to, and living in, the UK. We expect to award between 4 and 6 studentships to international candidates in 2024.
  • ARIES students benefit from bespoke graduate training and ARIES provides £2,500 to every student for access to external training, travel and conferences, on top of all Research Costs associated with the project. Excellent applicants from quantitative disciplines with limited experience in environmental sciences may be considered for an additional 3-month stipend to take advanced-level courses.
  • ARIES is committed to equality, diversity, widening participation and inclusion in all areas of its operation. We encourage enquiries and applications from all sections of the community regardless of gender, ethnicity, disability, age, sexual orientation and transgender status. Academic qualifications are considered alongside non-academic experience, and our recruitment process considers potential with the same weighting as past experience.
  • All ARIES studentships may be undertaken on a part-time or full-time basis, visa requirements notwithstanding.
  • For further information, please contact the supervisor. To apply for this Studentship follow the instructions at the bottom of the page or click the 'apply now' link.
  • ARIES is required by our funders to collect Equality and Diversity Information from all of our applicants. The information you provide will be used solely for monitoring and statistical purposes; it will remain confidential, and will be stored on the UEA sharepoint server. Data will not be shared with those involved in making decisions on the award of Studentships, and will have no influence on the success of your application. It will only be shared outside of this group in an anonymised and aggregated form. You will be ask to complete the form by the University to which you apply.
  • ARIES funding is subject to UKRI terms and conditions. Postgraduate Researchers are expected to live within reasonable distance of their host organisation for the duration of their studentship. See https://www.ukri.org/publications/terms-and-conditions-for-training-funding/ for more information

Applications open

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